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Evolution of Insects in terms of the Implicate and Explicate Orders.

Part IIId


The Blattaria fauna of the Kuznets Basin
( Siberia, Russia )




Overall geographic and geologic situation of the Kuznets Basin.

The (palaeo-)geographical situation of the Kuznets basin is indicated in the next map :

Figure 1 :  Palaeogeography and insect localities in the Upper Carboniferous. From the palaeogeographic maps of  TERMIER and TERMIER, SCHUCHERT, and BUBNOFF. The largest of the North American black dots :  Mazon Creek (Illinois).
(After HENNIG, 1969. New localities added by PONT and SCHLEE, 1981).


The units, here presented as 'etages' or 'stages' of the Kuzbas (basin of Kuznets), are no strictly chronostratigraphic units such as, for example, the Kazan stage, the Kungur stage, etc. They are so-called 'suites' of which the age is not yet precisely determined.
The occurring complexes of fossil insects do show, however, that these suites are indeed more or less parallel with the time-lines, because the disappearance and appearance of higher taxa in them do point to a time-effect [our theory cannot entirely sustain this, however]. Differences according to facies will at most express themselves in differences in lower taxa (families, genera). Data about the lithology of the basin and especially that of the insect localities are absolutely necessary for a correct chronostratigraphical evaluation of the relevant strata.

The basin of Kuznets geographically lies just North of the 'geometric' center of Asia :  North of western China / western Mongolia, and just east of the town Novosibirsk, and south-west of the town Krasnojarsk (but closer to Novosibirsk). Politically the Kuzbas lies entirely within the territory of the Sovjet Union (Russia).
In the far north of the basin there is the town Tomsk at the river Tomj. Upstream to the south, also at this river, we have the town Kemerovo, which is the administrative center of the Kuzbas. At about the center of the basin (more or less to the west) there is the town Leninsk-Kuznetsky, and in the southern part of the basin there is, at the river Tomj, the town Novokuznetsk (alias Stalinsk).
The kuzbas is a relatively low-lying region covered with taiga, through which the river Tomj runs, bordered in the west by the Salair-heights, in the east by the Kuznetsk Alataus, and far to the south lies the Altai.
The region is at least 300 km long and about 100 km wide, making it, as to its extent, comparable with that of Holland.
North of the Kuzbas there is the low-land river-basin of the river Obj, of which the Tomj is a tributary discharging itself into the Obj just a little north of Tomsk.
The heights around the Kuzbas consist of Praecambrian and lower and middle Palaeozoic strata.
The basin itself consists of a thick (8-10 km) coal-bearing formation, ranging from the Carboniferous until, and including, the lower Jurassic.
Quaternary ('anthropogenous') deposits normally lie on these strata, while Neogenous, Palaeogenous, and Cretacean continental deposits are only present along the outer edge of the mountains surrounding the coal-bearing strata.
The coal-measures beginning with the Ostrogsk-stage, lie on lime-stone and shales with marine Tournaisian and Visean fossils.
The Kuzbas coal-measures consist of clastic sediments, up to 10 km thick as has been said, and take up the central lower part of the basin.
The estimated ages of the respective stages (suites) are given in the Table below. Meanwhile it is found that the Kuznets-stage has upper Permian (Ufimian) age instead of lower Permian.

Geologic Age Suite Sub-suite Formation Thickness  in meters
Jurassic Conglomeratic -- -- 700-900
Lower Trias Mal'tsevian -- Upper Mal'tsevian (variegated)
Lower Mal'tsevian (tuffogenous)
300-400
280-300
Upper Permian Kol'chuginian Upper Kol'chuginian
(Yerunakovian)

Lower Kol'chuginian

Gramoteinian
Leninskian

Uskatian
Il'inskian

1200-1400
500-700
400-500
600-900
Upper Lower Permian Kuznyetskian -- -- 700-800
Lower Permian
and
Upper and Middle
Carboniferous
Balakhonkian Upper Balakhonkian
(Lower Permian)

Lower Balakhonkian
Middle and Upper
Carboniferous
Usyatskian
Kemerovian
Ishanovo-Intermediate

Alykayevian
Mazurovian

100-170
160-1200

200-600
300-550

Namurian Ostrogian -- -- 200-600
Visean -- -- -- --
Tournaisian -- -- -- --

Table 1.  Stratigraphy of the Coal-bearing Formation in Kuzbas
(Accepted at a stratigraphic conference in 1954) (From NALIVKIN, 1962 (1973))

See also next Table (further down) in which the order of ranking of the chronostratigraphic units are changed and where also the ages based on the entomofaunae are a little differently estimated.

Correlation with European strata is not yet (around 1973) definitively determined. Differences in fauna and flora are in this case for an important part accounted for by biogeographic and climatic differences instead of by differences of age.

The conditions under which the coal-bearing strata were formed were changing as time progressed (NALIVKIN, 1962 (1973)) (see next Table) :
The Ostrogsk-stage is (as a lithological body) deposited in near-shore conditions, first in a shallow sea, and thereafter on a near-coast swampy plain, resulting in the first coal-seams.
The Balachon-stages have been deposited entirely on land, whereby the sea was retreating several hundreds of kilometers from the Kuzbas.
The Koljtsjuginsk Series (= Kuznets-stage + Iljinsk-stage + Jerunakowsk-stage) was deposited on the vast river-plains of a large enclosed region separated from the sea by thousands of kilometers.

The Ostrogsk-stage has (here always :  within the Kuzbas) only a small number of thin coal-seams.
The Balachon-stages have 30-40 coal-seams of 'economic thickness' of which many are up to 10 meter thick. Some of them reach a thickness of 13-18 meter.
After this a (virtually) coal-free sequens follows, which corresponds with the Kuznets-stage and the Iljinsk stage (together 2200 m thick).
The Jerunakowsk-stage again has a large number of coal-seams of 'economic thickness', 0.5-7.25 m.
The Maljtsewsk-stage (Trias), following upon it, has no coal, but the Jurassic strata, lying on top of it, do have coal again.

The approximate palaeogeographic position of the Kuzbas we here indicate on a palaeogeographic small map taken from WINDLEY, 1986 (see also Figure 1, above) :

Figure 2 :  Pangea in Permian times.
(After WINDLEY, 1986)

The basin of Kuznets is part of the Angara-geosyncline. The latter constituted the southern (at the time perhaps eastern) mobile belt of the Siberian Kraton (Platform). [Far] to the west of this Kraton there was the Ural-geosyncline, and west of the latter there was the European Kraton (= Russian Platform).
Of the tectonic structures the Hercynic are the most important in the Kuzbas.
The folds in the Balachon-stages are, at the edges of the basin, more or less complex. They simplify towards the center of the basin.
In the Koljtsjuginsk Series the strata are relatively gently folded.

The next Table gives the same (sub)stages of the Kuzbas with a change in ranking order and some adjustments of their age based on the entomofauna, according to ROHDENDORF, et al., 1961.  The Jerunakowsk, the Iljinsk, and the Kuznetsk suites (stages) together form the Koljtsjuginsk Series.

Subdivision of deposits
(accepted by the stratigraphic conference in 1956)
Geologic Age
(according to ROHDENDORF, et al., 1961)
Suites (stages) Sub-suites (sub-stages)                                                     
Maljtsewsk -- T1  ( Lower Triassic)                 
Jerunakowsk Tailugansk
Gramotjeinsk
Leninsk
P2  ( Upper Permian)                 
Iljinsk Uskatsk
Kazankowo-markinsk
P2  ( Upper Permian)                 
Kuznetsk -- P1  ( Lower Permian)                 
Upper Balachon Usjatsk
Kemerowsk
Isjanowsk
Promjezjutotsjnaja
C3  ( Upper Carboniferous)         
Lower Balachon Alikajewsk
Mazurowsk
C2  ( Middle Carboniferous)         
Ostrogsk -- C1  ( Lower Carboniferous)         

Table 2.  Stratigraphic subdivision of the Palaeozoic of the coal-bearing formation of the basin of Kuznets.
(After ROHDENDORF, et al., 1961)

The Kuznets-stage nowadays (ROHDENDORF et RASNITSIN, 1980 assessed to be of upper Permian age (P2) (ufimian)) is not subdivided into sub-stages. Biostratigraphically it is, according to MARTYNOVA in ROHDENDORF, et al., 1961, clearly divisible into three horizonts.
A precise determination of the geologic age of the stages and sub-stages is (MARTYNOVA, ibidem) not yet possible. In the above Table these ages are given (as preliminary directives) as based on the fossil insects.

As to this "based on the fossil insects" we must note the following :
As a consequence of our noëtic theory of evolution, the determination of the age of a given geological formation or deposit (stratum), and its correlation with other geological formations or deposits elsewhere, is not possible on the basis of biostratigraphy because in the latter it is assumed that organismic species evolve in time into other species resulting in a genealogic tree of them ascending through time from early to later epochs. In our theory it is, however, assumed that succession of different organismic species is not the result of material descent from each other. We hold that such a succession of species, as they appear one after the other in the Explicate Order (and evident by the fossil record) is wholly dictated by prevailing ecological conditions. The latter are existential conditions of strategy-contents immaterially produced in the Implicate Order. These strategy-contents are not produced from one another but each one is independently constructed in that Order. If species are produced from one another at all, this always is a production via the Implicate Order, and the point in time, as seen from the Explicate Order, of their appearance in that Order, totally depends on the presence at that time of the proper existental conditions in the Explicate Order. And so it may happen that a 'primitive' species appears in the latter Order much later in time than its more 'derived' cousin, because only at that (later) time the existential conditions for it came into being. See for all this the
  Theoretic Intermezzo  following upon Part VII.  This Intermezzo (and its sequel) extensively deals with our noëtic theory of evolution (so far developed). For the leading role of ecological conditions, see (apart from what we already wrote about it in earler documents) in organic evolution (the successive apearance, in the Explicate Order, of different organismic species), especially about halfway in the (first document on the mentioned) Theororetic intermezzo from the Section dealing with  Organic strategy-contents and their existential conditions in the Explicate Orderonwards (including the Sequel of the Intermezzo).
So the only way to determine the relative age of a geologic formation or deposit is its position within the overall series of formations or deposits of which it is a part. If, and only if, such a series has not been too much deformed by tectonics (i.e. intensively folded, contorted, or having been warped upside down), a given layer or deposit is older than the one on top of it. And the lithological structure of a pack of layers may, in cases, indicate a gap in time between two such layers.
And of course all this holds for sedimentary rocks only.



We shall now present the fossil  Blattida (cockroaches) found (in the form of impressions of tegmina only) in the Palaeozoic of the basin of Kuznets. ["middle Carboniferous" and "upper Carboniferous" in the Table below, may, perhaps, respectively be "middle Upper-Carboniferous" and "upper Upper-Carboniferous"]

In most of the next images the archedictyon will not be given
(as it was done in the present figure) although it is present.


Phyloblatta  carbonica BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen about 36.5 mm.
A - tegmen (forewing),  B, C -- archedictyon.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Balachonoblatta  zheltojarica BECKER-MIGDISOVA.
Family Archimylacridae.
Length of fragment about 26.3 mm.
Length tegmen probably 30 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Hemimylacrella  leptophlebiopsis BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen about 39 mm. Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Sibiroblatta  flexuosa BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 25 mm.
Probable length of tegmen about 28.5 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)
Grypoblattites  flabellatus BECKER-MIGDISOVA
Family Archimylacridae (or maybe Poroblattinidae).
Length of impression 20 mm.
Probable length of tegmen 25-26 mm.
Locality :  Zjelty Jar.
Middle Carboniferous of Kuznets Basin, Siberia, Russia.
(After ROHDENDORF et al., 1961).

Branches of CuA tending to descend steeper onto the wing's hind margin.


Grypoblattites  tortuosus BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 21.3 mm.
Probable length of tegmen about 28-30 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Archaeotiphites  petroblattinoides BECKER-MIGDISOVA.
Family Archimylacridae (or maybe Poroblattinidae, despite its relatively large size).
Length of impression 26.4 mm.
Probable length of tegmen about 29 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Archaeotiphites  captiosus BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 24.9 mm.
Probable length of tegmen about 28 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  lineata BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 28.2 mm.
Probable length of tegmen about 32-34 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  martynovi BECKER-MIGDISOVA.
Family Archimylacridae.
Probable length of tegmen about 26 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  curviramosa BECKER-MIGDISOVA.
Family Archimylacridae.
Probable length of tegmen 25-27 mm.
Locality :  Porivaisk mine, Basin of Kuznets, Siberia, Russia.
upper Balachon stage. Upper Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  distincta MARTYNOV.
Family Archimylacridae.
Length (variable) of tegmen 26-34 mm. Ten specimens found.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  alykaevensis BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 32 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  curvinervosa BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 32 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  biramosa BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 31 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  triangularis BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 25-30 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  adjuncta BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 30 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  propinqua BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 27 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  kuznetskiensis BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 29 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  visenda BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 30 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  incelebrata BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 28.8 mm. Probable length of tegmen 35 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  tomiensis BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 30 mm. Probable length of tegmen 36-37 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  semiplena BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 34 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)


Phyloblatta  sibirica  M.ZALESSKY.
Family Archimylacridae.
Length of tegmen 27-34 mm.
( Four specimens found in Zjelty Jar, and one in Tserimitskino)
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Lower image :  The specimen found in Tserimitskino (same stage).


Phyloblatta  foveolata BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 23.1 mm. Probable length of tegmen 28 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  indistincta BECKER-MIGDISOVA.
Family Archimylacridae.
Probable length of tegmen about 28 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  dubia BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 29.2 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  aliena BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 30 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  arealata BECKER-MIGDISOVA.
Family Archimylacridae.
Probable length of tegmen about 40 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  multinervosa BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 22 mm. Probable length of tegmen about 30-40 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  khalfini BECKER-MIGDISOVA.
Family Archimylacridae.
Length of impression 29 mm. Probable length of tegmen about 31 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  furcatella MARTYNOV.
( Tomioblatta  furcatella MARTYNOV, 1931, p.79 Family Archimylacridae.
Length of impression 26 mm. Length of tegmen about 26.5 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta sp. I.
Family Archimylacridae.
Length of impression 28.2 mm. Length of tegmen about 26.5 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta sp. II.
Family Archimylacridae.
Length of impression 33.3 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Miaroblatta  balachonensis BECKER-MIGDISOVA
Family Archimylacridae.
Length of impression 28.5-31.0 mm. (two specimens found)
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Ignaroblatta  panda BECKER-MIGDISOVA
Family Archimylacridae.
Length of impression 20.3 mm. Probable length of tegmen 25 mm
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Ignaroblatta  sibirica BECKER-MIGDISOVA
Family Archimylacridae.
Length of impression 30 mm. Probable length of tegmen 34-35 mm
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Ignaroblatta  rectisubcosta BECKER-MIGDISOVA
Family Archimylacridae.
Length of impression 22.9 mm.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Ignaroblatta sp.
Family Archimylacridae.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Aphthoroblattina  irregularis BECKER-MIGDISOVA.
Length of impression 18 mm. Probable length of tegmen 25-30 mm.
Family Archimylacridae.
Locality :  Zjelty Jar, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Phyloblatta  picturata BECKER-MIGDISOVA.
Length of tegmen about 18 mm.
Family Archimylacridae.
Locality :  Zavjalowo, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)

Relatively small form.


Phyloblatta  paulitransversinervis BECKER-MIGDISOVA.
Length of impression 16 mm. Probable length of tegmen 25 mm.
Family Archimylacridae.
Locality :  Zavjalowo, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)


Sysciophlebia  zavjalovensis BECKER-MIGDISOVA.
Length of impression 23.9 mm. Probable length of tegmen 24.5 mm.
Family Spiloblattinidae.
Locality :  Zavjalowo, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)


Zavjaloblatta  rotundata BECKER-MIGDISOVA.
Probable length of tegmen about 20 mm.
Family Spiloblattinidae.
Locality :  Zavjalowo, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage, Alikajewsk sub-stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)


Caenoblatta  angaridensis  M.ZALESSKY.
Family Archimylacridae.
Locality :  Tsjerimitschkino, Basin of Kuznets, Siberia, Russia.
Lower Balachon stage.
Middle Carboniferous.
(After ROHDENDORF, et al., 1961)


Phyloblatta  poryvaikaensis  BECKER-MIGDISOVA.
Family Archimylacridae.
Probable length of tegmen 44 mm.
Locality :  Porivaisk mine, Basin of Kuznets, Siberia, Russia.
Upper Balachon stage. Isjanowskaja sub-stage.
Upper Carboniferous.
(After ROHDENDORF, et al., 1961)


Phyloblatta  lata  BECKER-MIGDISOVA.
Family Archimylacridae.
Probable length of tegmen 40 mm.
Locality :  Porivaisk mine, Basin of Kuznets, Siberia, Russia.
Upper Balachon stage. Isjanowskaja sub-stage.
Upper Carboniferous.
(After ROHDENDORF, et al., 1961)


Phyloblatta  mala  BECKER-MIGDISOVA.
Family Archimylacridae.
Length of tegmen 27 mm.
Locality :  Porivaisk mine, Basin of Kuznets, Siberia, Russia.
Upper Balachon stage. Isjanowskaja sub-stage.
Upper Carboniferous.
(After ROHDENDORF, et al., 1961)


Olethroblatta  tomiensis  BECKER-MIGDISOVA.
Family Spiloblattinidae.
Length of impression 22.1 mm.
Locality :  Porivaisk mine, Basin of Kuznets, Siberia, Russia.
Upper Balachon stage. Isjanowskaja sub-stage.
Upper Carboniferous.
(After ROHDENDORF, et al., 1961)


Metaxyblatta  gromotuchensis  BECKER-MIGDISOVA.
Family Mylacridae.
Length of impression 18.5 mm. Probable length of tegmen about 24 mm.
Locality :  Porivaisk mine, Basin of Kuznets, Siberia, Russia.
Upper Balachon stage. Isjanowskaja sub-stage.
Upper Carboniferous.
(After ROHDENDORF, et al., 1961)


Ishanoblattina (Drepanoblattina)  fodinensis  BECKER-MIGDISOVA.
Family Poroblattinidae.
Length of impression 22 mm. Probable length of tegmen about 24-25 mm.
Locality :  Porivaisk mine, Basin of Kuznets, Siberia, Russia.
Upper Balachon stage. Isjanowskaja sub-stage.
Upper Carboniferous.
(After ROHDENDORF, et al., 1961)

* * *

These, then, were the fossil Blattida found (up until 1961) in the palaeozoic strata of the basin of Kuznets.
Four families have been identified, viz., Archimylacridae, Spiloblattinidae, Mylacridae, and Poroblattinidae. The majority of them is formed by the Archimylacridae.




(Will be continued . . . )


With this we conclude the exposition of blattarian fauna of the Kuznetz Basin. In the next document we will describe a new wing-type, elytropterygia (shield-wingedness).



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